• value of the temptation to defect (T ).
By investigating the effect of these parameters, we
aim to explore the impact of the link update rules and
to investigate the evolution of cooperation when ab-
stainers are present in the population.
Although some work has considered coevolv-
ing link weights when considering the Prisoner’s
Dilemma in a spatial environment, to our knowledge
the investigation of an Optional Prisoner’s Dilemma
game on a spatial environment, where both the strate-
gies and link weights are evolved, has not been stud-
ied to date.
The results show that cooperation emerges even
in extremely adverse scenarios where the temptation
of defection is almost at its maximum. It can be ob-
served that the presence of the abstainers are funda-
mental in protecting cooperators from invasion. In
general, it is shown that, when the coevolutionary
rules are used, cooperators do much better, being also
able to dominate the whole population in many cases.
Moreover, for some settings, we also observe inter-
esting phenomena of cyclic competition between the
three strategies, in which abstainers invade defectors,
defectors invade cooperators and cooperators invade
abstainers.
The paper outline is as follows: Section 2 presents
a brief overview of the previous work in both spa-
tial evolutionary game theory with dynamic networks
and in the Optional Prisoner’s Dilemma game. Sec-
tion 3 gives an overview of the methodology em-
ployed, outlining the Optional Prisoner’s Dilemma
payoff matrix, the coevolutionary model used (Monte
Carlo simulation), the strategy and link weight update
rules, and the parameter values that are varied in order
to explore the effect of coevolving both strategies and
link weights. Section 4 features the results. Finally,
Section 5 summarizes the main conclusions and out-
lines future work.
2 RELATED WORK
The use of coevolutionary rules constitute a new trend
in evolutionary game theory. These rules were first in-
troduced by Zimmermann et al. (2001), who proposed
a model in which agents can adapt their neighbour-
hood during a dynamical evolution of game strategy
and graph topology. Their model uses computer sim-
ulations to implement two rules: firstly, agents play-
ing the Prisoner’s Dilemma game update their strat-
egy (cooperate or defect) by imitating the strategy of
an agent in their neighbourhood with a higher pay-
off; and secondly, the network is updated by allowing
defectors to break their connection with other defec-
tors and replace the connection with a connection to
a new neighbour selected randomly from the whole
network. Results show that such an adaptation of the
network is responsible for an increase in cooperation.
In fact, as stated by Perc and Szolnoki (2010),
the spatial coevolutionary game is a natural upgrade
of the traditional spatial evolutionary game initially
proposed by Nowak and May (1992), who consid-
ered static and unweighted networks in which each
individual can interact only with its immediate neigh-
bours. In general, it has been shown that coevolving
the spatial structure can promote the emergence of co-
operation in many scenarios (Wang et al., 2014; Cao
et al., 2011), but the understanding of cooperative be-
haviour is still one of the central issues in evolutionary
game theory.
Szolnoki and Perc (2009) proposed a study of the
impact of coevolutionary rules on the spatial version
of three different games, i.e., the Prisoner’s Dilemma,
the Snow Drift and the Stag Hunt game. They intro-
duce the concept of a teaching activity, which quanti-
fies the ability of each agent to enforce its strategy on
the opponent. It means that agents with higher teach-
ing activity are more likely to reproduce than those
with a low teaching activity. Differing from previ-
ous research (Zimmermann et al., 2004, 2001), they
also consider coevolution affecting either only the de-
fectors or only the cooperators. They discuss that, in
both cases and irrespective of the applied game, their
coevolutionary model is much more beneficial to the
cooperators than that of the traditional model.
Huang et al. (2015) present a new model for the
coevolution of game strategy and link weight. They
consider a population of 100 × 100 agents arranged
on a regular lattice network which is evolved through
a Monte Carlo simulation. An agent’s interaction is
described by the classical Prisoner’s Dilemma with a
normalized payoff matrix. A new parameter, ∆/δ, is
defined as the link weight amplitude and is calculated
as the ratio of ∆/δ. They found that some values of
∆/δ can provide the best environment for the evolu-
tion of cooperation. They also found that their coevo-
lutionary model can promote cooperation efficiently
even when the temptation of defection is high.
In addition to investigations of the classical Pris-
oner’s Dilemma on spatial environments, some exten-
sions of this game have also been explored as a means
to favour the emergence of cooperative behaviour.
For instance, the Optional Prisoner’s Dilemma game,
which introduces the concept of abstinence, has been
studied since Batali and Kitcher (1995). In their
work, they proposed the opt-out or “loner’s” strategy
in which agents could choose to abstain from play-
ing the game, as a third option, in order to avoid co-
The Optional Prisoner’s Dilemma in a Spatial Environment: Coevolving Game Strategy and Link Weights
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